The answer is Human beings.
According to the Bible, sin entered the world through human beings. In the Book of Genesis, Adam and Eve, the first humans, disobeyed God by eating the forbidden fruit from the Tree of Knowledge of Good and Evil. This act of disobedience is considered the original sin, and it is through this act that sin and its consequences, such as death and suffering, entered the world (Genesis 3:1-19).
Natural selection is a scientific concept related to the process of evolution and is not associated with the theological concept of sin in the Bible. Therefore, it is not considered responsible for sin and its consequences according to biblical teachings.
While God created the world and established the moral laws, the Bible does not attribute the responsibility for sin to God. Instead, it emphasizes human free will and the choice to disobey God's commandments as the source of sin.
In the Bible, angels are spiritual beings created by God. While some angels, such as Satan, rebelled against God, the responsibility for sin entering the human world is attributed to human actions, specifically the disobedience of Adam and Eve.